Summer-green. Rhizome stout, ascending to erect, covered with persistent stipe bases; scales thin, rusty-brown, ovate-lanceolate, 2-3 mm long 0.5 mm broad. Stipe much shorter than lamina, tufted, slender, stramineous sometimes purplish, 3-20 cm long; lamina narrowly lanceolate, deeply bipinnatifid to bipinnate (basal pinnules free), 10-20 cm long, 3-6 cm wide; all except upper pairs of pinnae reflexed, 2-4 cm long, sessile; pinnules cut nearly to costa, obtuse, dentate, adaxial basal pinnule enlarged, herbaceous, glabrous, fertile even in small leaves; ultimate segments oblong to elliptic, serrate or sharply lobed at margin; very short spines present on the adaxial side of costae. Sori medial or basal on lower veinlets; indusia oblong, J-shaped or reniform; margins somewhat erose.
Distribution: Taiwan, Japan, Shikoku and Kyushu (Yakushima), and S. Korea.
Athyrium Roth 蹄蓋蕨屬
Pseudocystopteris Ching 假冷蕨屬
Terrestrial ferns; rhizome usually stout and erect, rarely long creeping; scales thin, lanceolate or ovate and light brown, some narrow and black . Stipe green, stramineous, or purple, with 2 vascular strands at base and uniting into U-shaped strand at middle rachies; lamina pinnate to decompound; veins all free; rachis and costae grooved and connected on dorsal side, some species with spines on the dorsal side of rachies, costae and costulate. Sori oblong or round, usually crossing a veinlet, some species with Diplazium type. Indusia linear, J-shape or reniform. About 24 species in Taiwan, most species in Taiwan grown at altitude 1000 to 3000 m.